Introduction
Species of the genus Nilotonia are knovvn from standing vvaters, streams, seepage areas, and vvarm springs, and are reported from vvidely scattered areas in Africa, vvith a more limited range in Europe (Cook, 1974; Viets, 1978).
Anatolia (Turkey), a land bridge betvveen Asia and Europe, has an important zoogeographic, ecological, and geologic position. To date, 5 species belonging to different genera of the subfamily Nilotoniinae have been reported from Anatolia (Özkan, 1982; Özkan and Bader, 1988; Soysal and Özkan, 1989; Özkan and Soysal, 1989; Özkan and Boyacı, 1990).
The subgenus Manotonia of the genus Nilotonia differs from other subgenera in the absence of elongated, thin setae on IV-leg; (if elongated setae are present, they are strong and broad) vvithout elongated terminal or subterminal seta on IV-leg/6; lateral setae on IV-leg/6 typically reduced in number (5-0) and short;
sclerotization on dorsum and ventrum variable; dorsal seta on P1 typically absent.
According to Bader (1980), the genus Nilotonia includes 6 subgenera: Dartiella Viets, 1929, Nilotonia Thor, 1905, Tadjikodartia Bader, 1980; Dartia Soar, 1917; Dartonia Viets, 1929; and Parvadartia Bader, 1980.
Later, the subgenera Mamersonia Viets, 1954, Manotonia Viets, 1935, and Guanacastonia and Bolivartonia (Goldschmidt, 2004) were assigned to the genus Nilotonia (Panesar, 2004).
During the survey of the freshwater fauna of the West Taurus Mountains (Anatolia), N. (M.) tegulata was collected in a high mountain brook.


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Materials and Methods
The material was collected among mosses on stones and washed under pressured tap water. The water mites Dissection of body parts and orientation of specimens on slides were done before the specimens were identified.
Taxonomy
Family: Anisitsiellidae
Genus: Nilotonia, Thor, 190S
Nilotonia (Manotonia) tegulata (Viets, 1951)
(Figures 1-2)
Material Examined: Turkey, East Anatolia, Taurus Mountains; 20.05.2000, 299, 2dd; 26.08.2003, 499, 2dd, 1200 m. The specimens were collected from mosses on stones in a small spring, close to Köprüçay stream anddeposited in S.D. University, Fisheries Faculty. Eğirdir, İsparta.

Descriptiorı
Male
Dorsal side: Body with secondary sclerotization and posteriorly setose, 588/521 um in size. Dorsal shield covers more than one half of body fused with postantenniformae platelets; integument finely lineated; dorsal shield anteriorly with 2 projections at anterior, 334/234 nm in size.
Gnathosoma: Capitulum with a short rostrum, 153 um in length; chelicerae 328 um, cheliceral claw 70 um in length; P4 with 4 distoventral setae; dorsal and ventral lengths of palpal segments: 25-80-47-100-25 = 277 pm, 23-55-33-83-23 = 217 um, respectively.
Figüre 1. Nilotonia tegulata; Male: A) dorsal view, B) ventral view, Female: C) ventral view, D) dorsal view.

Figure 2. Nilotonia tegulata; Male: A-D) legs i, II, III, IV, E) chelicera, F) gnathosoma.

Ventral side: First coxal groups 163 um long, second coxal group 221 um in length; posterior region to fourth coxae with secondary sclerotization and setose. Anterior coxal groups not fused medially; suture line betvveen third and fourth coxae incomplete; glands on the third coxae present; genital field extending anteriorly to posterior margin of fourth coxae, 125 um in length; genital area with a well pre- and post-genital sclerite, which are fused with the ventral shield.
Legs: Svvimming hairs absent; fourth leg vvithout claws but terminating in 4 short setae; claws on other
legs with 2 clavvlets; legs of male not exhibiting sexual dimorphism. I-, II-, III-leg segments bear a number of long and slender setae. Lengths of leg segments: leg I 50-62-55-67-98-115 = 447 um, leg II: 55-65-60-85-118-142 = 525 um, leg III; 60-68-90-140-143-134 = 635 um, leg IV: 82-62-88-115-147-147 = 641 um.
Female: Body narrovved anteriorly and 508/429 um in size; unsclerotized parts of integument finely lineated; each eye with 2 lenses; postantenniformae platelets 66 um in length; dorsal plate 154/158 um in size; there are 2 small platelets anteriorly of postocularia platelets; capitulum with a short rostrum, 292 um in length; chelicerae 328 um, cheliceral claw 90 um in length; P2 with a seta in ventrolateral position and P4 bears 4 setae on 2 small ventral projections located distoventrally; palp delicate and with a few setae. Dorsal and ventral length of palpal segments; 25-100-55-122-35 = 337 pm, 27-73-33-102-33 = 268 pm, respectively. First coxal groups 152 pm, second coxal group 198 pm in length and with secondary sclerotization. Anterior coxal groups not fused medially; suture üne betvveen third and fourth coxae incomplete; genital field anteriorly extending betvveen the fourth coxae and 113 pm in length; svvimming hairs absent; fourth leg vvithout clavvs but terminating in 2 short setae; clavvs on other legs vvith 2 clavvlets; legs bear a large number of stocky setae. Lengths of leg segments; leg I: 50-67-55-83-93-113 = 461 pm, leg II: 55-83-65-63-115-127 = 508 pm, leg III: 70-80-73-150-138-138 = 649 pm, leg IV: 70-78-85-135-157-165 = 690 pm.

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Key to the subgenera of Nilotonia (Thor, 1905)
1. The medial margins of the third and forth coxae parallel to each other anteriorly to genital field .
- Third coxae vvith pointed medial margin anteriorly to the genital field.............4
2. IV-leg/5 distally vvith svvimming hairs; dorsum vvith 3 platelets of moderate size.....Nilotonia
3. IV-leg/3-5 vvith clearly elongated thin setae, but vvithout svvimming setae; most of the species dorsally vvith extensive secondary sclerotization .
.........................Mamersonia
- If large setae on segment IV-leg/3-5 are present distally, then they are not thin; IV-leg/6 vvith only a fevv small lateral setae and never vvith elongated terminal seta; dorsum vvith posteromedial plate of either enlarged or moderate size . . . .Manotonia
4. First coxae extending posteriorly, typically nearly reaching the anterior border of the genital field and IV-leg/6 vvith a strong terminal seta......
Not so
.5
5. Coxaglandularia IV protruding together vvith parts
of Coxa3 over posterior parts of Coxa2.......
.........................Tadjikodartia
6. Gap between first and second acetabula longer than length of first acetabula, and basal sclerites of acetabula with very visible chitinization; chitinization of posterior 2 pairs typically fused on each side; acetabula reduced in size; sexual dimorphism leads to typically shaped terminal
segments in the second pair legs of males .....
............................. Dartia
- Gap between first and second acetabula shorter than length of first acetabula, but typically acetabula extend along the medial border of the genital flaps; if the acetabula are reduced in size, then the basal sclerites are not, or only partly chitinized ..................... Dartiella

Results and Discussion
In the revision of the family Anisitsiellidae Koenike by Panesar (2004), the following species were included in the subgenus Manotonia: Nilotonia gracilipalpis Lundblad 1942 (Africa, Ethiopia); N. musicola (Walter, 1935) (Africa, Ivory Coast), N. shivai Panesar, 2004 (Asia, India), N. petri Cook, 1966 (Africa, Cameroon), N. scutata Cook, 1966 (Africa, Cameroon), N. tegulata (Viets, 1951) (Africa, Algeria; Europe, Italy), N. testudinata(Cook, 1966) (Africa, Liberia), and N. violacea Lundblad, 1951 (Africa, Kenya). Recently, Pesic and Saboori (2006) described N. (Manotonia) persica from Iran.
Most species of the subgenus are known from Africa; only N. (Manotonia) tegulata has been reported from Africa as well as from European parts of the Mediterranean region.
N.(Manotonia) tegulatadiffers from all of the African species of the subgenus Manotonia, in the absence of a complete ventral plate. Moreover, it is easily distinguished from similar species by the presence of a large dorsal plate that covers one-third of the dorsum and is fused with the postantenniformae platelets; the body has posteriorly secondary sclerotization, which is setose, while leg IV terminates in 4 short setae in the male, 2 of these shorter than the other 2 setae; genital area with well developed pre- and post-genital sclerites fused with the ventral plate; swimming hairs absent; ventral of P2 not denticulate; anterior coxal groups not fused medially; capitulum with a narrow and blunt rostrum and not bearing a projecting part in dorso-lateral position (Viets, 1916; Lundblad, 1952, 1956; Viets, 1956; Bader, 1980; Gerecke, 1991).
Gerecke (1991) pointed out that there are no characters to separate the female of N. (Manotonia) tegulatafrom other sclerotized species of Nilotonia.
In the accompanying water mite fauna Sperchon rostratus Lundblad, Trichothyas petrophila (Michael), Atractides longirostris Walter, Bandakia concreta Thor, Brachypoda mutila (Walter), Monatractides adoratus Di Sabatino and Gerecke, M. stadleri(Walter), M. lusitanicus (Lundblad), and an unidentified species of the genus Lebertiawere observed.
Before we can give an interpretation in terms of systematical problems in the genus Nilotonia, further information is needed on the fauna of the neighboring geographical areas, as well as on the phylogenetic relationships between the genera and species of the family.

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